Scrat's epic pursuit of his elusive acorn catapults him outside of Earth, where he accidentally sets off a series of cosmic events that transform and threaten the planet. To save themselves from peril, Manny, Sid, Diego, and the rest of the herd leave their home and embark on a quest full of thrills and spills, highs and lows, laughter and adventure while traveling to exotic new lands and encountering a host of colorful new characters.
Jennifer Lopez....Shira (voice)
Melissa Rauch....Francine (voice)
Simon Pegg....Buck (voice)
John Leguizamo....Sid (voice)
Stephanie Beatriz....Gertie (voice)
Nick Offerman....Gavin (voice)
Max Greenfield....Roger (voice)
Seann William Scott....Crash (voice)
Josh Peck....Eddie (voice)
Keke Palmer....Peaches (voice)
Denis Leary....Diego (voice)
Queen Latifah....Ellie (voice)
Jesse Tyler Ferguson....Shangri Llama (voice)
Ray Romano....Manny (voice)
Filmography links and data courtesy of The Internet Movie Database.